Detailed study of the Khatyspyt Formation in arctic Siberia has shown that, contrary to common expectations, the earliest ichnofabric is of late Ediacaran age, records a food-seeking behavior, and is found in a distal carbonate ramp setting where it co-occurs with fossils of Avalon-type Ediacaran soft-bodied organisms. Preserved depth of bioturbation reaches 5 cm. The ichnofabric solely comprises Nenoxites, the oldest known meniscate trace fossil, which is interpreted as burrows with terminal backfill structure formed as a result of active displacement of sediment to form a tunnel within the substrate, followed by emplacement of the material behind the animal as it progressed through the sediment.
